make_parameters {photosynthesis} | R Documentation |
Make lists of parameters for photosynthesis
Description
Make lists of parameters for photosynthesis
make_leafpar
make_enviropar
make_bakepar
make_constants
Usage
make_leafpar(replace = NULL, use_tealeaves)
make_enviropar(replace = NULL, use_tealeaves)
make_bakepar(replace = NULL)
make_constants(replace = NULL, use_tealeaves)
Arguments
replace |
A named list of parameters to replace defaults.
If |
use_tealeaves |
Logical. Should leaf energy balance be used to calculate leaf temperature (T_leaf)? If TRUE, |
Details
Constants:
Symbol | R | Description | Units | Default |
| D_c0 | diffusion coefficient for CO2 in air at 0 °C | m / s | |
| D_h0 | diffusion coefficient for heat in air at 0 °C | m / s | |
| D_m0 | diffusion coefficient for momentum in air at 0 °C | m / s | |
| D_w0 | diffusion coefficient for water vapor in air at 0 °C | m / s | |
| epsilon | ratio of water to air molar masses | none | 0.622 |
| G | gravitational acceleration | m / s | 9.8 |
| eT | exponent for temperature dependence of diffusion | none | 1.75 |
| R | ideal gas constant | J / mol / K | 8.31 |
| sigma | Stephan-Boltzmann constant | W / m / K | |
| f_sh | function to calculate constant(s) for Sherwood number | none | NA |
| f_nu | function to calculate constant(s) for Nusselt number | none | NA |
Baking (i.e. temperature response) parameters:
Symbol | R | Description | Units | Default |
| Ds_gmc | empirical temperature response parameter | J / mol / K | 487 |
| Ds_Jmax | empirical temperature response parameter | J / mol / K | 388 |
| Ea_gammastar | empirical temperature response parameter | J / mol | 24500 |
| Ea_gmc | empirical temperature response parameter | J / mol | 68900 |
| Ea_Jmax | empirical temperature response parameter | J / mol | 56100 |
| Ea_KC | empirical temperature response parameter | J / mol | 81000 |
| Ea_KO | empirical temperature response parameter | J / mol | 23700 |
| Ea_Rd | empirical temperature response parameter | J / mol | 40400 |
| Ea_Vcmax | empirical temperature response parameter | J / mol | 52200 |
| Ea_Vtpu | empirical temperature response parameter | J / mol | 52200 |
| Ed_gmc | empirical temperature response parameter | J / mol | 149000 |
| Ed_Jmax | empirical temperature response parameter | J / mol | 121000 |
Environment parameters:
Symbol | R | Description | Units | Default |
| C_air | atmospheric CO2 concentration | umol/mol | 420 |
| O | atmospheric O2 concentration | mol/mol | 0.21 |
| P | atmospheric pressure | kPa | 101 |
| PPFD | photosynthetic photon flux density | umol / m / s | 1500 |
| RH | relative humidity | none | 0.5 |
| wind | windspeed | m / s | 2 |
Leaf parameters:
Symbol | R | Description | Units | Default |
| leafsize | leaf characteristic dimension | m | 0.1 |
| gamma_star | chloroplastic CO2 compensation point (T_leaf) | umol/mol | NA |
| gamma_star25 | chloroplastic CO2 compensation point (25 °C) | umol/mol | 37.9 |
| g_mc | mesophyll conductance to CO2 (T_leaf) | mol / m / s | NA |
| g_mc25 | mesophyll conductance to CO2 (25 °C) | mol / m / s | 0.4 |
| g_sc | stomatal conductance to CO2 | mol / m / s | 0.4 |
| g_uc | cuticular conductance to CO2 | mol / m / s | 0.01 |
| J_max25 | potential electron transport (25 °C) | umol / m / s | 200 |
| J_max | potential electron transport (T_leaf) | umol / m / s | NA |
| k_mc | partition of g_mc to lower mesophyll | none | 1 |
| k_sc | partition of g_sc to lower surface | none | 1 |
| k_uc | partition of g_uc to lower surface | none | 1 |
| K_C25 | Michaelis constant for carboxylation (25 °C) | umol / mol | 268 |
| K_C | Michaelis constant for carboxylation (T_leaf) | umol / mol | NA |
| K_O25 | Michaelis constant for oxygenation (25 °C) | umol / mol | 165000 |
| K_O | Michaelis constant for oxygenation (T_leaf) | umol / mol | NA |
| phi_J | initial slope of the response of J to PPFD | none | 0.331 |
| R_d25 | nonphotorespiratory CO2 release (25 °C) | umol / m / s | 2 |
| R_d | nonphotorespiratory CO2 release (T_leaf) | umol / m / s | NA |
| theta_J | curvature factor for light-response curve | none | 0.825 |
| T_leaf | leaf temperature | K | 298 |
| V_cmax25 | maximum rate of carboxylation (25 °C) | umol / m / s | 150 |
| V_cmax | maximum rate of carboxylation (T_leaf) | umol / m / s | NA |
| V_tpu25 | rate of triose phosphate utilization (25 °C) | umol / m / s | 200 |
| V_tpu | rate of triose phosphate utilisation (T_leaf) | umol / m / s | NA |
If use_tealeaves = TRUE
, additional parameters are:
Constants:
Symbol | R | Description | Units | Default |
| c_p | heat capacity of air | J / g / K | 1.01 |
| R_air | specific gas constant for dry air | J / kg / K | 287 |
Baking (i.e. temperature response) parameters:
Symbol | R | Description | Units | Default |
Environment parameters:
Symbol | R | Description | Units | Default |
| E_q | energy per mole quanta | kJ / mol | 220 |
| f_par | fraction of incoming shortwave radiation that is photosynthetically active radiation (PAR) | none | 0.5 |
| r | reflectance for shortwave irradiance (albedo) | none | 0.2 |
| T_air | air temperature | K | 298 |
| T_sky | sky temperature | K | NA |
Leaf parameters:
Symbol | R | Description | Units | Default |
| abs_l | absorbtivity of longwave radiation (4 - 80 um) | none | 0.97 |
| abs_s | absorbtivity of shortwave radiation (0.3 - 4 um) | none | 0.5 |
| g_sw | stomatal conductance to H2O | mol / m / s | NA |
| g_uw | cuticular conductance to H2O | mol / m / s | NA |
| logit_sr | stomatal ratio (logit transformed) | none | NA |
Optional leaf parameters:
Symbol | R | Description | Units | Default |
| delta_ias_lower | effective distance through lower internal airspace | um | NA |
| delta_ias_upper | effective distance through upper internal airspace | um | NA |
| A_mes_A | mesophyll surface area per unit leaf area | none | NA |
| g_liqc25 | liquid-phase conductance to CO2 (25 °C) | mol / m / s | NA |
| g_liqc | liquid-phase conductance to CO2 (T_leaf) | mol / m / s | NA |
| g_iasc_lower | internal airspace conductance to CO2 in lower part of leaf (T_leaf) | mol / m / s | NA |
| g_iasc_upper | internal airspace conductance to CO2 in upper part of leaf (T_leaf) | mol / m / s | NA |
Value
make_leafpar
: An object inheriting from class leaf_par()
make_enviropar
: An object inheriting from class enviro_par()
make_bakepar
: An object inheriting from class bake_par()
make_constants
: An object inheriting from class constants()
References
Buckley TN and Diaz-Espejo A. 2015. Partitioning changes in photosynthetic rate into contributions from different variables. Plant, Cell & Environment 38: 1200-11.
Examples
bake_par = make_bakepar()
constants = make_constants(use_tealeaves = FALSE)
enviro_par = make_enviropar(use_tealeaves = FALSE)
leaf_par = make_leafpar(use_tealeaves = FALSE)
leaf_par = make_leafpar(
replace = list(
g_sc = set_units(0.3, mol / m^2 / s),
V_cmax25 = set_units(100, umol / m^2 / s)
), use_tealeaves = FALSE
)