make_parameters {photosynthesis}R Documentation

Make lists of parameters for photosynthesis

Description

Make lists of parameters for photosynthesis

make_leafpar

make_enviropar

make_bakepar

make_constants

Usage

make_leafpar(replace = NULL, use_tealeaves)

make_enviropar(replace = NULL, use_tealeaves)

make_bakepar(replace = NULL)

make_constants(replace = NULL, use_tealeaves)

Arguments

replace

A named list of parameters to replace defaults. If NULL, defaults will be used.

use_tealeaves

Logical. Should leaf energy balance be used to calculate leaf temperature (T_leaf)? If TRUE, tleaf() calculates T_leaf. If FALSE, user-defined T_leaf is used. Additional parameters and constants are required, see make_parameters().

Details

Constants:

Symbol R Description Units Default
Dc,0D_{\mathrm{c},0} D_c0 diffusion coefficient for CO2 in air at 0 °C m2^2 / s 1.29×1051.29\times 10^{-5}
Dh,0D_{\mathrm{h},0} D_h0 diffusion coefficient for heat in air at 0 °C m2^2 / s 1.90×1051.90\times 10^{-5}
Dm,0D_{\mathrm{m},0} D_m0 diffusion coefficient for momentum in air at 0 °C m2^2 / s 1.33×1051.33\times 10^{-5}
Dw,0D_{\mathrm{w},0} D_w0 diffusion coefficient for water vapor in air at 0 °C m2^2 / s 2.12×1052.12\times 10^{-5}
ϵ\epsilon epsilon ratio of water to air molar masses none 0.622
GG G gravitational acceleration m / s2^2 9.8
eTeT eT exponent for temperature dependence of diffusion none 1.75
RR R ideal gas constant J / mol / K 8.31
σ\sigma sigma Stephan-Boltzmann constant W / m2^2 / K4^4 5.67×1085.67\times 10^{-8}
fShf_\mathrm{Sh} f_sh function to calculate constant(s) for Sherwood number none NA
fNuf_\mathrm{Nu} f_nu function to calculate constant(s) for Nusselt number none NA

Baking (i.e. temperature response) parameters:

Symbol R Description Units Default
Ds,gmcD_\mathrm{s,gmc} Ds_gmc empirical temperature response parameter J / mol / K 487
Ds,JmaxD_\mathrm{s,Jmax} Ds_Jmax empirical temperature response parameter J / mol / K 388
Ea,ΓE_\mathrm{a,\Gamma *} Ea_gammastar empirical temperature response parameter J / mol 24500
Ea,gmcE_\mathrm{a,gmc} Ea_gmc empirical temperature response parameter J / mol 68900
Ea,JmaxE_\mathrm{a,Jmax} Ea_Jmax empirical temperature response parameter J / mol 56100
Ea,KCE_\mathrm{a,KC} Ea_KC empirical temperature response parameter J / mol 81000
Ea,KOE_\mathrm{a,KO} Ea_KO empirical temperature response parameter J / mol 23700
Ea,RdE_\mathrm{a,Rd} Ea_Rd empirical temperature response parameter J / mol 40400
Ea,VcmaxE_\mathrm{a,Vcmax} Ea_Vcmax empirical temperature response parameter J / mol 52200
Ea,VtpuE_\mathrm{a,Vtpu} Ea_Vtpu empirical temperature response parameter J / mol 52200
Ed,gmcE_\mathrm{d,gmc} Ed_gmc empirical temperature response parameter J / mol 149000
Ed,JmaxE_\mathrm{d,Jmax} Ed_Jmax empirical temperature response parameter J / mol 121000

Environment parameters:

Symbol R Description Units Default
CairC_\mathrm{air} C_air atmospheric CO2 concentration umol/mol 420
OO O atmospheric O2 concentration mol/mol 0.21
PP P atmospheric pressure kPa 101
PPFD\mathrm{PPFD} PPFD photosynthetic photon flux density umol / m2^2 / s 1500
RH\mathrm{RH} RH relative humidity none 0.5
uu wind windspeed m / s 2

Leaf parameters:

Symbol R Description Units Default
dd leafsize leaf characteristic dimension m 0.1
Γ\Gamma* gamma_star chloroplastic CO2 compensation point (T_leaf) umol/mol NA
Γ25\Gamma*_{25} gamma_star25 chloroplastic CO2 compensation point (25 °C) umol/mol 37.9
gmcg_\mathrm{mc} g_mc mesophyll conductance to CO2 (T_leaf) mol / m2^2 / s NA
gmc,25g_\mathrm{mc,25} g_mc25 mesophyll conductance to CO2 (25 °C) mol / m2^2 / s 0.4
gscg_\mathrm{sc} g_sc stomatal conductance to CO2 mol / m2^2 / s 0.4
gucg_\mathrm{uc} g_uc cuticular conductance to CO2 mol / m2^2 / s 0.01
Jmax,25J_\mathrm{max,25} J_max25 potential electron transport (25 °C) umol / m2^2 / s 200
JmaxJ_\mathrm{max} J_max potential electron transport (T_leaf) umol / m2^2 / s NA
kmck_\mathrm{mc} k_mc partition of g_mc to lower mesophyll none 1
ksck_\mathrm{sc} k_sc partition of g_sc to lower surface none 1
kuck_\mathrm{uc} k_uc partition of g_uc to lower surface none 1
KC,25K_\mathrm{C,25} K_C25 Michaelis constant for carboxylation (25 °C) umol / mol 268
KCK_\mathrm{C} K_C Michaelis constant for carboxylation (T_leaf) umol / mol NA
KO,25K_\mathrm{O,25} K_O25 Michaelis constant for oxygenation (25 °C) umol / mol 165000
KOK_\mathrm{O} K_O Michaelis constant for oxygenation (T_leaf) umol / mol NA
ϕJ\phi_J phi_J initial slope of the response of J to PPFD none 0.331
Rd,25R_\mathrm{d,25} R_d25 nonphotorespiratory CO2 release (25 °C) umol / m2^2 / s 2
RdR_\mathrm{d} R_d nonphotorespiratory CO2 release (T_leaf) umol / m2^2 / s NA
θJ\theta_J theta_J curvature factor for light-response curve none 0.825
TleafT_\mathrm{leaf} T_leaf leaf temperature K 298
Vc,max,25V_\mathrm{c,max,25} V_cmax25 maximum rate of carboxylation (25 °C) umol / m2^2 / s 150
Vc,maxV_\mathrm{c,max} V_cmax maximum rate of carboxylation (T_leaf) umol / m2^2 / s NA
Vtpu,25V_\mathrm{tpu,25} V_tpu25 rate of triose phosphate utilization (25 °C) umol / m2^2 / s 200
VtpuV_\mathrm{tpu} V_tpu rate of triose phosphate utilisation (T_leaf) umol / m2^2 / s NA

If use_tealeaves = TRUE, additional parameters are:

Constants:

Symbol R Description Units Default
cpc_p c_p heat capacity of air J / g / K 1.01
RairR_\mathrm{air} R_air specific gas constant for dry air J / kg / K 287

Baking (i.e. temperature response) parameters:

Symbol R Description Units Default

Environment parameters:

Symbol R Description Units Default
EqE_q E_q energy per mole quanta kJ / mol 220
fPARf_\mathrm{PAR} f_par fraction of incoming shortwave radiation that is photosynthetically active radiation (PAR) none 0.5
rr r reflectance for shortwave irradiance (albedo) none 0.2
TairT_\mathrm{air} T_air air temperature K 298
TskyT_\mathrm{sky} T_sky sky temperature K NA

Leaf parameters:

Symbol R Description Units Default
αl\alpha_\mathrm{l} abs_l absorbtivity of longwave radiation (4 - 80 um) none 0.97
αs\alpha_\mathrm{s} abs_s absorbtivity of shortwave radiation (0.3 - 4 um) none 0.5
gswg_\mathrm{sw} g_sw stomatal conductance to H2O mol / m2^2 / s NA
guwg_\mathrm{uw} g_uw cuticular conductance to H2O mol / m2^2 / s NA
logit(sr)\mathrm{logit}(sr) logit_sr stomatal ratio (logit transformed) none NA

Optional leaf parameters:

Symbol R Description Units Default
δias,lower\delta_\mathrm{ias,lower} delta_ias_lower effective distance through lower internal airspace um NA
δias,upper\delta_\mathrm{ias,upper} delta_ias_upper effective distance through upper internal airspace um NA
Ames/AA_\mathrm{mes} / A A_mes_A mesophyll surface area per unit leaf area none NA
gliq,c,25g_\mathrm{liq,c,25} g_liqc25 liquid-phase conductance to CO2 (25 °C) mol / m2^2 / s NA
gliq,cg_\mathrm{liq,c} g_liqc liquid-phase conductance to CO2 (T_leaf) mol / m2^2 / s NA
gias,c,lowerg_\mathrm{ias,c,lower} g_iasc_lower internal airspace conductance to CO2 in lower part of leaf (T_leaf) mol / m2^2 / s NA
gias,c,upperg_\mathrm{ias,c,upper} g_iasc_upper internal airspace conductance to CO2 in upper part of leaf (T_leaf) mol / m2^2 / s NA

Value

make_leafpar: An object inheriting from class leaf_par()
make_enviropar: An object inheriting from class enviro_par()
make_bakepar: An object inheriting from class bake_par()
make_constants: An object inheriting from class constants()

References

Buckley TN and Diaz-Espejo A. 2015. Partitioning changes in photosynthetic rate into contributions from different variables. Plant, Cell & Environment 38: 1200-11.

Examples

bake_par = make_bakepar()
constants = make_constants(use_tealeaves = FALSE)
enviro_par = make_enviropar(use_tealeaves = FALSE)
leaf_par = make_leafpar(use_tealeaves = FALSE)

leaf_par = make_leafpar(
  replace = list(
    g_sc = set_units(0.3, mol / m^2 / s),
    V_cmax25 = set_units(100, umol / m^2 / s)
  ), use_tealeaves = FALSE
)

[Package photosynthesis version 2.1.4 Index]