make.geosse.t {diversitree} | R Documentation |

Prepare to run time dependent GeoSSE (Geographic State Speciation and Extinction) on a phylogenetic tree and character distribution. This function creates a likelihood function that can be used in maximum likelihood or Bayesian inference.

```
make.geosse.t(tree, states, functions, sampling.f=NULL, strict=TRUE,
control=list(), truncate=FALSE, spline.data=NULL)
```

`tree` |
A phylogenetic tree, in |

`states` |
A vector of character states, each of which must be 0
(in both regions/widespread; AB), 1 or 2 (endemic to one region; A or
B), or |

`functions` |
A named character vector of functions of time. See details. |

`sampling.f` |
Vector of length 3 with the estimated proportion of
extant species in states 0, 1 and 2 that are included in the
phylogeny. A value of |

`strict` |
The |

`control` |
List of control parameters for the ODE solver. See
details in |

`truncate` |
Logical, indicating if functions should be truncated to zero when negative (rather than causing an error). May be scalar (applying to all functions) or a vector (of length 7). |

`spline.data` |
List of data for spline-based time functions. See
details in |

.

Please see `make.bisse.t`

for further details.

`make.geosse.t`

returns a function of class `geosse.t`

.

The `funtions`

is a vector of named functions of time. For
example, to have speciation rates be linear functions of time, while
the extinction and dispersal rates be constant with respect to time,
one can do

functions=rep(c("linear.t", "constant.t"), c(3, 4))

. The functions here must have `t`

as their first
argument, interpreted as time back from the present. See
`make.bisse.t`

for more information, and for some
potentially useful time functions.

The function has argument list (and default values):

f(pars, condition.surv=FALSE, root=ROOT.OBS, root.p=NULL, intermediates=FALSE)

The parameter vector pars is ordered `sA`

, `sB`

, `sAB`

,
`xA`

, `xB`

, `dA`

, `dB`

. Unresolved clade methods
are not available for GeoSSE. With three states, it would rapidly
become computationally infeasible. The arguments of this function are
also explained in `make.bisse`

.

`starting.point.geosse`

produces a first-guess set of parameters,
ignoring character states.

This computer intensive code is experimental!

Jonathan Rolland

FitzJohn R.G. 2012. Diversitree: comparative phylogenetic analyses of diversification in R. Methods in Ecology and Evolution. 3, 1084-1092.

FitzJohn R.G., Maddison W.P., and Otto S.P. 2009. Estimating trait-dependent speciation and extinction rates from incompletely resolved phylogenies. Syst. Biol. 58:595-611.

Goldberg E.E., Lancaster L.T., and Ree R.H. 2011. Phylogenetic inference of reciprocal effects between geographic range evolution and diversification. Syst. Biol. 60:451-465.

Maddison W.P., Midford P.E., and Otto S.P. 2007. Estimating a binary character's effect on speciation and extinction. Syst. Biol. 56:701-710.

Nee S., May R.M., and Harvey P.H. 1994. The reconstructed evolutionary process. Philos. Trans. R. Soc. Lond. B Biol. Sci. 344:305-311.

`constrain`

for making submodels and reduce number
of parameters, `find.mle`

for ML parameter estimation,
`mcmc`

for MCMC integration, `make.bisse`

and
`make.bisse.t`

for further relevant examples.

The help page for `find.mle`

has further examples of ML
searches on full and constrained BiSSE models. Things work similarly
for GeoSSE and GeoSSE.t, just with different parameters.

See `make.geosse`

for explanation of the base model.

```
## Due to a change in sample() behaviour in newer R it is necessary to
## use an older algorithm to replicate the previous examples
if (getRversion() >= "3.6.0") {
RNGkind(sample.kind = "Rounding")
}
## Parameter values
pars <- c(1.5, 0.5, 1.0, 0.7, 0.7, 2.5, 0.5)
names(pars) <- diversitree:::default.argnames.geosse()
## Simulate a tree
set.seed(5)
phy <- tree.geosse(pars, max.t=4, x0=0)
## See the data
statecols <- c("AB"="purple", "A"="blue", "B"="red")
plot(phy, tip.color=statecols[phy$tip.state+1], cex=0.5)
## Create your list of functions. Its length corresponds to the number
## of parameters (speciation, extinction and dispersal) you want to
## estimate.
## For an unconstrained model, at least 7 parameters are estimated for
## sA, sB, sAB, xA, xB, dA, dB.
## In the case you want to define a model with linear functions of
## speciation and extinction, and constant dispersal:
functions <- rep(c("linear.t", "constant.t"), c(5, 2))
## Create the likelihood function
lik <- make.geosse.t(phy, phy$tip.state, functions)
## This function will estimate a likelihood from 12 parameters.
argnames(lik)
## Imagine that you want to get an estimate of the likelihood from a
## known set of parameters.
pars <- c(0.01,0.001,0.01,0.001,0.01,0.001,0.02,0.002,0.02,0.002,0.1,0.1)
names(pars)<-argnames(lik)
lik(pars) # -640.1644
## A guess at a starting point from character independent birth-death
## model (constant across time) .
p <- starting.point.geosse(phy)
#it only gives 7 parameters for time-constant model.
names(p)
## it can be modified for time-dependent with a guess on the slopes of
## speciation and extinction rates.
p.t<-c(p[1],0.001,p[2],0.001,p[3],0.001,p[4],0.001,p[5],0.001,p[6],p[7])
names(p.t)<-argnames(lik)
## Start an ML search from this point (takes from one minute to a very
## long time depending on your computer).
## Not run:
fit <- find.mle(lik, p.t, method="subplex")
fit$logLik
coef(fit)
## End(Not run)
## A model with constraints on the dispersal rates.
lik.d <- constrain(lik, dA ~ dB)
##Now dA and dB are the same parameter dB.
argnames(lik.d)
##The parameter dA must be removed from maximum likelihood initial parameters
## Not run:
fit.d <- find.mle(lik.d, p.t[-which(names(p.t)=="dA")])
fit$logLik
coef(fit)
## End(Not run)
```

[Package *diversitree* version 0.9-16 Index]